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Pleuroweisia schliephackei (Pleuroweisia schliephackei)

Description

Plants growing in turf or mats, green to yellow-brown above, light to dark brown below. Stems seldom branching, to 1-3 cm in length, transverse section oval to rounded-triangular or pentagonal, central strand present, strong, outer cortex with smaller lumens, usually thick-walled, hyalodermis absent; axillary hairs of 3-10 cells, all hyaline or basal 1-2 brownish and with thicker walls; weakly radiculose or with red-brown tomentum. Leaves often distant or crowded, occasionally secund, appressed-incurved, often twisted when dry, spreading when moist, ligulate to lanceolate, occasionally triangular or acuminate, 1.0-1.5-2.0 mm in length, upper lamina strongly keeled, deeply grooved along costa, margins plane or weakly recurved in lower 1/2, entire, occasionally finely crenulate or weakly denticulate above; apex broadly obtuse to sharply acute, usually apiculate, occasionally acuminate or somewhat cucullate; base scarcely differentiated in shape or ovate; costa ending at the apiculus or sometimes a fewcells below apex or becoming short-excurrent as a mucro, seldom stoutly excurrent, superficial cells elongate or occasionally short-rectangular to quadrate near apex ventrally, elongate dorsally and generally papillose, 2-3 cells across costa ventrally at midleaf, costal transverse section oval to reniform, stereid band single, strong, semicircular to oval, epidermis absent ventrally (the guide cells superficially exposed) to weakly developed, epidermis usually distinct dorsally, guide cells 2-4 in 1 layer, hydroid strand absent; upper laminal cells subquadrate, occasionally hexagonal or elongate transversely or longitudinally in patches, 5-7-9-15-m in width, 1-2:1-2, walls thin to greatly thickened, superficially flat to bulging, occasionally in extraplanar rows (very seldom bistratose); papillae multifid, often massively so, centered over the lumens or simple to bifid, scattered, rarely absent; basal cells differentiated in a small group at base of costa, short-rectangular, little wider than upper cells, 2-4:1, walls usually thick-walled. Dioicous. Perichaetia terminal on short lateral branches, inner leaves ovate-acuminate, 1.0-1.5 mm in length, convolute-sheathing, lower cells short-rhomboidal to near apex. Perigonia lateral. Seta 0.3-0.8 cm in length, 1 per perichaetium, yellow-brown, twisted clockwise below, occasionally counterclockwise above; theca 0.5-1.0-1.5mm in length, yellow-brown to brown, ovoid to elliptical, exothecial cells rectangular, walls thin, stomates phaneropore, at base of theca, annulus of two rows of weakly vesiculose cells; peristome absent. Operculum long-rostrate, 0.4-0.6-1.8 mm in length, sometimes longer than the theca, inclined, cells in straight rows. Calyptra cucullate, smooth, 1.2-1.5-2.0 mm in length. Spores 9-12-19 -m in diameter, light brown, weakly to strongly papillose. Laminal KOH color reaction yellow to yellow-orange. Reported chromosome number n = 13.A rather large but singularly homogeneous group distributed mainly in tropical, arctic or mountane areas of the world. Anoectangium is unusual in that the distinctions between the many species are mainly in characters considered variable in other genera. A close study of the Middle American representation (Zander 1977c) resulted in extensive synonymy with only one species, A. aestivum (Pl. 38, f. 10-13), recognized for the area. The most common expression of the species in the area was found to have small, superficially bulging laminal cells with multifid papillae centered over the lumens. Considered weakly distinguishable were certain uncommon, mostly local variants with various leaf shapes and usually superficially flat, larger upper laminal cells having thickened walls, often transversely or longitudinally elongate medially, the laminal papillae low, simple, seldom multifid. Some collections, however, showed independent segregation of these features.

Taxonomic tree

  • Domain: Eukarya

    • Kingdom: Plantae

      • Phylum:

        • Class: Bryopsida

          • Order: Pottiales

            • Family: Pottiaceae

              • Genus: Pleuroweisia